U.S. Secretary of State John Kerry laid out what he called overwhelming evidence of Russian complicity in the shooting down of Malaysia Airlines flight MH17 as international horror deepened over the fate of the victims' remains.
Kerry demanded that Moscow take responsibility for actions of pro-Russian rebels in eastern Ukraine whom Washington suspects of downing the jet with a missile, and expressed disgust at their "grotesque" mishandling of the bodies.
Television images of the rebel-held crash sites, where the remains of victims had lain decomposing in fields among their personal belongings, have turned initial shock and sorrow after Thursday's disaster into anger.
Emotions ran high in the Netherlands, the home country of about two thirds of the 298 people who died in the Boeing 777. The Dutch foreign minister has said the nation is "furious" to hear bodies were being "dragged around", while relatives and church leaders demanded they be rapidly returned home.
However, the departure of dozens of corpses loaded into refrigerated railway wagons was delayed on Sunday as Ukrainian officials and rebels traded blame over why the train had not yet left the war zone, and where or when international investigators would be able to check it.
In Washington, Kerry criticized Russian President Vladimir Putin and threatened "additional steps" against Moscow.
"Drunken separatists have been piling bodies into trucks and removing them from the site," he said on NBC television on Sunday. "What's happening is really grotesque and it is contrary to everything President Putin and Russia said they would do."
Moscow denies any involvement in the disaster and has blamed the Ukrainian military. While stopping short of direct blame on Moscow, Kerry put forward the most detailed U.S. accusations so far that Russia provided the insurgents with the sophisticated anti-aircraft systems used to down the aircraft.
British Foreign Secretary Philip Hammond echoed the criticism, urging Moscow to ensure international investigators had access to the crash sites. "Russia risks becoming a pariah state if it does not behave properly," he told Sky television.
After lying for two days in the summer heat, the bodies had been removed from much of the crash site by Sunday, leaving only bloodstained military stretchers along the side of the road.
Emergency workers, who have to navigate reporting both to the authorities in Kiev and the rebels who control the crash site and other areas in the Donetsk region, will now need to pick through the debris spread across the Ukrainian steppe.
As Ukraine accused the rebels of hiding evidence relating to the loss of the airliner, a separatist leader said items thought to be the stricken jet's "black boxes" were now in rebel hands.
Kerry said the United States had seen supplies moving into Ukraine from Russia in the last month, including a 150-vehicle convoy of armored personnel carriers, tanks and rocket launchers given to the separatists.
It had also intercepted conversations about the transfer to separatists of the Russian radar-guided SA-11 missile system which it blames for the Boeing 777's destruction. "It's pretty clear that this is a system that was transferred from Russia," Kerry said in an interview on CNN.
"There's enormous amount of evidence, even more evidence that I just documented, that points to the involvement of Russia in providing these systems, training the people on them," he said on CBS.
The disaster has sharply deepened the Ukrainian crisis in which the separatists in the Russian-speaking east have been fighting government forces since protesters in Kiev forced out a pro-Moscow president and Russia annexed Crimea in March.
The United States has already imposed sanctions on individuals and businesses close to Putin but Kerry indicated that President Barack Obama might go further. "The president is prepared to take additional steps," he told Fox News, although he ruled out sending in U.S. troops.