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The ax is falling again at Wells Fargo's mortgage origination unit, as refinancing activity continues to slow.
The bank on Wednesday sent 60-day notices of displacement to 1,800 employees across the country, citing a slowdown in activity throughout 2012 and early 2013.
"After evaluating the current market and our business needs, we are responding to this change in demand and to better align and increase the efficiency of our organization," said Wells Fargo spokesperson Kate Pulley in a statement.
It's unclear how much Wells Fargo—the nation's largest mortgage provider—needs to rightsize to adjust to the new normal. The bank has already cut 3,000 jobs in origination and servicing, but activity has been bouncing around.
A brief drop in mortgage rates over Labor Day weekend caused refinancing activity to pick back up. News this week that the Federal Reserve would maintain its pace of bond buying sent yields lower, which in effect will cause mortgage rates to fall.
(Read more: Mortgages: To lock or not to lock?)
The average rate on a 30-year fixed mortgage for the week ended Sept. 13 was 4.5 percent, versus 3.53 percent in May, according to the Mortgage Bankers Association.
With the mortgage boom officially over, banks are closing facilities and units that have been servicing customers looking to refinance their home loans.
Other Wall Street banks are making similar moves, as a sharp rise in rates has kept consumers from taking out or refinancing mortgages.
JPMorgan laid off more than 2,000 employees in early August—about half of them in originations, according to a person familiar with the situation. The bank had said in February that it would cut 17,000 jobs (largely in the mortgage servicing unit, which handles bad loans) by the end of 2014.
The August round of layoffs represented the first time the bank had moved to downsize its origination business, which surged as mortgage rates went to historic lows.
In August, Bank of America notified 2,100 employees that their jobs were being cut; Wells Fargo has laid off more than 3,000 since July. Citigroup confirmed in early September the July closure of an office in Danville, Ill., that affected 120 jobs.
"The Danville facility was originally established to handle the surge in demand for refinancing," said Citi spokesman Mark Rodgers, who added that other telephone sales positions supporting mortgage banking would be cut.
Consumers rushed to take advantage of low rates to refinance mortgages. Though the Federal Reserve is expected to keep short-term rates low for some time, talk of potentially slowing the Fed's stimulus program has led government bond yields—to which mortgages are tied—straight up.
While most industry executives expected the market to slow as interest rates rose, few expected it to screech to a halt as it has.
(Read more: Map: Tracking the recovery)
At an investor conference this week, JPMorgan Chase CFO Marianne Lake called the drop-off in mortgage activity "dramatic and rapid."
Executives expected that an improving economy and rising housing prices would spur more people to take out mortgages to buy homes, but Lake said that volume won't compensate for the refinancing business that has dried up.
According to the MBA, mortgage applications fell 13.5 percent in the week ended Sept. 6, and the portion of mortgages from refinancing fell only slightly. William J. Rogers, CEO of Atlanta-based SunTrust, said his company's mortgage applications were down 40 percent in August and July.
(Read more: Can the mortgage market crash again?)
Just a month shy of third-quarter earnings, bank caution is contagious. Both Lake and Tim Sloan, Wells Fargo's CFO, suggested that their mortgage volumes would fall more than 30 percent. Sloan projected Wells Fargo's originations at just $80 billion in the third quarter, down from $112 billion last quarter, with few areas for the bank to make up that business.
In a low-rate environment, an expanding mortgage business was a bank's saving grace. In the third quarter, it could be the industry's Achilles' heel.
—By CNBC's Kayla Tausche. Follow her on Twitter: @KaylaTausche