The occupation of the refuge, about six hours' drive from Portland, began on Jan. 2. Armed militants demanded the release of two local ranchers who were imprisoned on arson charges for burning public lands. They also called for federal lands that had been in private hands, generations ago, to be turned over to ranchers or to local government.
The standoff appeared to be faltering in late January, when several prominent occupiers — including Ammon and Ryan Bundy — were arrested while venturing out of the refuge. A spokesman for the group, LaVoy Finicum, was killed, and several of the remaining occupiers heeded calls by the Bundys to go home.
But four of them held out for another two weeks, before revealing in a live-streamed conversation Wednesday that they intended to surrender on Thursday. They invoked the death of Mr. Finicum as evidence that the government did not want a peaceful conclusion, saying that they feared being killed, too.
On Thursday morning, Ms. Fiore urged the last four militants to surrender so they could continue to spread their message. "A dead man can't talk, a dead man can't write," she told them. "We have to just stay together, stay alive."
Sean Anderson, his wife, Sandy, and Jeff Banta emerged after 9:30 a.m. But Mr. Fry, still on the phone with Mr. Seim, Ms. Hall and an F.B.I. agent, resisted until almost 11 a.m.
"I'm paying taxes, and it's going for abortion," Mr. Fry said, and also for "murder of millions in the Middle East." Later, he said, "My concern is that if I go to prison, I'll probably be raped."
In a telephone interview, Mr. Fry's father, William Fry Jr., 56, said he had spoken with his son after the arrest. "He said, 'Hi, I'm O.K., I'm going to be fine.' "
The two had been in contact during the occupation, and the elder Mr. Fry said he supported his son in "trying to make a change, to save our country from the problems that we've got."
About 50 or 60 cars were parked at the roadblock outside the sanctuary on Thursday, where protest sympathizers mixed with journalists. Thomas Wagner, 32, an unemployed security guard from Christmas Valley, Ore., stood atop his pickup truck in full military fatigues, and said, "I came here to support these four patriots, to let them know that they are not being abandoned."
In Burns, the town closest to the refuge, people raised American flags up and down the main street to celebrate the occupation's end. "This is better than the Fourth of July," said Bekka Riess, 15, beaming as she put flags up. "Maybe now we can finally get our town back."